The lyrics reference American Revolutionary Patrick Henry's quote "give me liberty or give me death" with the line "liberty or death, what we so proudly hail". The lyric, "To secure peace, is to prepare for war" refers to the Latin adage Si vis pacem, para bellum ("If you want peace, prepare [for] a war"). The lyrics containing rattlesnake imagery are inspired by Benjamin Franklin's essay suggesting the rattlesnake is a good symbol for the American Spirit.

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The song was never performed live until the European 2012 tour celebrating the 20th anniversary of the Black Album, starting at the tour's first show in Prague. At the end of the concert, Hetfield mentioned that the band liked the live sound of songs ("Don't Tread on Me" and "My Friend of Misery") that they played live for the first time. Previously, Hetfield was said to have disliked the song, as revealed in an April 2001 interview with Playboy magazine, in which he stated: "There are some songs on there I don't like. 'Don't Tread on Me', probably not one of my favorite songs musically."[6]

"Don't Tread on Me" is a song by American heavy metal band Metallica from their self-titled fifth album. The title is connected with the American Revolutionary War. The words "Don't Tread on Me" constitute the motto of the Gadsden flag, and the snake image on the flag is pictured on the cover of the album.

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The Broncos were barely competitive during their 10-year run in the AFL and their first seven years in the NFL. They did not complete a winning season until 1973. In 1977, four years later, they qualified for the playoffs for the first time in franchise history and advanced to Super Bowl XII. Since 1975, the Broncos have become one of the NFL's more successful teams, having suffered only six losing seasons. They have won eight AFC Championships and three Super Bowl championships (1997, 1998  2015 (50); sharing the record for second most Super Bowl appearances with the Dallas Cowboys, and Pittsburgh Steelers, right behind the New England Patriots record nine Super Bowl appearances.

The tune was named by journalist John J. Miller of the conservative news magazine National Review as one of the "50 Greatest Conservative Rock songs"; placed as No. 9 on the list, Miller praised it as a "tribute to the doctrine of peace through strength".[7]

The Denver Broncos are an American football team based in Denver, Colorado. The Broncos compete in the National Football League (NFL) as a member club of the league's American Football Conference (AFC) West division. The team began play in 1960 as a charter member of the American Football League and joined the NFL as part of the merger in 1970. The Broncos are owned by the Pat Bowlen trust. The Broncos have played at Sports Authority Field at Mile High since 2001, after previously playing at Mile High Stadium from 1960 to 2000.

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The main guitar riff has been sampled on the song "Surface Patterns" by Canadian industrial band Front Line Assembly, from their 1994 album Millennium.[4][5]

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Hetfield also said "Don't Tread on Me, I love the song, but it shocked a lot of people, because everyone thought it was pro-war when they thought we were anti-war, and alls we're doing is writing songs, we're not standing politically on any side. Don't Tread on Me was just one of those 'don't fuck with us' songs, and obviously referencing the flag and the snake and what it meant, that all tied into the black album and the snake icon on the album cover, and I think it's great to play that song live. We're over here in Europe playing it, and people aren't appalled by the songs. We haven't played it in Iraq or Iran yet, though."[3]

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The instrumental introduction uses an eight-bar phrase from "America", a popular song from the musical West Side Story. It is in a moderate tempo of 104 bpm in 128.[1]

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In 2019, Dee Gallant of Duncan, British Columbia, Canada, was walking her dog on a logging road near the town when she realized a cougar was approaching them. After shouting did not make it go away, she selected "Don't Tread on Me" as the loudest song on her cellphone, and when she played it, the cougar left.[8][9] The following week, after news of the event had spread, she received a friendly phone call from lead singer James Hetfield.[9][10]

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Hetfield said the song was a reaction to the anti-establishment tone of their album ...And Justice for All: "This is the other side of that. America is a fucking good place. I definitely think that. And that feeling came about from touring a lot. You find out what you like about certain places and you find out why you live in America, even with all the bad fucked-up shit. It's still the most happening place to hang out."[2]